Balance at

December

31, 2019

Additions

Deductions

Balance at

December 31, 2020

Accounts receivable and contract assets:

Billed receivables from collaboration partners

$

20,000

$

6,644

$

26,637

$

7

Unbilled receivables from collaboration partners

158

7,725

5,559

2,324

Total accounts receivable and contract assets

$

20,158

$

14,369

$

32,196

$

2,331

Contract liabilities:

Deferred revenue - Incyte

$

8,378

$

335

$

3,348

$

5,365

Deferred revenue - GBT

20,000

746

4,025

16,721

Total contract liabilities

$

28,378

$

1,081

$

7,373

$

22,086