Balance at

December

31, 2020

Additions

Deductions

Balance at December 31, 2021

Accounts receivable and contract assets:

Billed receivables from collaboration partners

$

7

$

10,931

$

(10,938

)

$

—

Unbilled receivables from collaboration partners

2,324

11,410

(10,755

)

2,979

Total accounts receivable and contract assets

$

2,331

$

22,341

$

(21,693

)

$

2,979

Contract liabilities:

Deferred revenue - Incyte

$

5,365

$

—

$

(4,097

)

$

1,268

Deferred revenue - GBT

16,721

174

(7,982

)

8,913

Total contract liabilities

$

22,086

$

174

$

(12,079

)

$

10,181