Balance at

December

31, 2021

Additions

Deductions

Balance at March 31, 2022

Accounts receivable and contract assets:

Billed receivables from collaboration partners

$

—

$

2,857

$

(2,857

)

$

—

Unbilled receivables from collaboration partners

2,979

3,181

(2,978

)

3,182

Total accounts receivable and contract assets

$

2,979

$

6,038

$

(5,835

)

$

3,182

Contract liabilities:

Deferred revenue - Incyte

$

1,268

$

—

$

(421

)

$

847

Deferred revenue - GBT

8,913

—

(1,987

)

6,926

Total contract liabilities

$

10,181

$

—

$

(2,408

)

$

7,773