Quarterly report pursuant to Section 13 or 15(d)

Commitments and Contingencies - Operating Leases (Details)

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Commitments and Contingencies - Operating Leases (Details)
$ in Thousands
 1 Months Ended 9 Months Ended
Jan. 08, 2019
ft²
Jan. 01, 2019
USD ($)
Mar. 31, 2015
ft²
Sep. 30, 2019
USD ($)
Operating lease, right-of-use asset       $ 16,496
Operating lease liability       20,403
Operating lease liability, long term   $ 100   18,425
Operating lease liability, short term       1,978
Accounting Standards Update 2016-02        
Operating lease, right-of-use asset   1,500   15,800
Operating lease liability   2,200    
Operating lease liability, long term   1,100   15,800
Operating lease liability, short term   $ 1,100    
Net of tenant allowances       $ 9,300
Lease operating, discounted borrowing rate       9.30%
Lease practical expedient   true    
Accounting Standards Update 2016-02 | Operating Lease, January 2019        
Operating lease liability, long term       $ 18,300
Operating lease liability, short term       700
Operating Lease March2015        
Area of office and laboratory space leased | ft² | ft²     21,488  
Operating lease liability, short term       $ 1,300
Operating Lease, January 2019        
Area of office and laboratory space leased | ft² | ft² 52,859      
Term of option to extend the lease       10 years
Letter of credit outstanding       $ 3,100