Balance at

December

31, 2020

Additions

Deductions

Balance at

March 31, 2021

Accounts receivable and contract assets:

Billed receivables from collaboration partners

$

7

$

2,397

$

(2,404

)

$

—

Unbilled receivables from collaboration partners

2,324

2,622

(2,324

)

2,622

Total accounts receivable and contract assets

$

2,331

$

5,019

$

(4,728

)

$

2,622

Contract liabilities:

Deferred revenue - Incyte

$

5,365

$

—

$

(809

)

$

4,556

Deferred revenue - GBT

16,721

71

(1,396

)

15,396

Total contract liabilities

$

22,086

$

71

$

(2,205

)

$

19,952