Quarterly report pursuant to Section 13 or 15(d)

Commitments and Contingencies - Operating Leases (Details)

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Commitments and Contingencies - Operating Leases (Details)
$ in Thousands
 1 Months Ended 3 Months Ended
Jan. 08, 2019
ft²
Jan. 01, 2019
Mar. 31, 2015
ft²
Mar. 31, 2020
USD ($)
Dec. 31, 2019
USD ($)
Operating lease, right-of-use asset       $ 15,294 $ 15,821
Operating lease liability, short term       1,287 1,037
Operating lease liability, long term       24,244 $ 24,018
Accounting Standards Update 2016-02          
Operating lease, right-of-use asset       15,800  
Operating lease liability, long term       15,800  
Net of tenant allowances expected to be received       $ 9,300  
Lease operating, discounted borrowing rate       9.30%  
Lease practical expedient   true      
Operating Lease, March 2015          
Area of office and laboratory space leased | ft² | ft²     21,488    
Operating lease termination       As of December 31, 2019, the 2015 Lease had terminated and all obligations relating the 2015 Lease were satisfied in full with no remaining balances as of that date  
Operating Lease, January 2019          
Area of office and laboratory space leased | ft² | ft² 52,859        
Term of option to extend the lease       10 years  
Letter of credit outstanding       $ 3,100  
Operating Lease, January 2019 | Accounting Standards Update 2016-02          
Operating lease liability, short term       1,300  
Operating lease liability, long term       $ 24,200