Balance at

December

31, 2020

Additions

Deductions

Balance at
September 30,
 2021

Accounts receivable and contract assets:

Billed receivables from collaboration partners

$

7

$

7,270

$

(7,277

)

$

—

Unbilled receivables from collaboration partners

2,324

8,431

(7,123

)

3,632

Total accounts receivable and contract assets

$

2,331

$

15,701

$

(14,400

)

$

3,632

Contract liabilities:

Deferred revenue - Incyte

$

5,365

$

—

$

(2,766

)

$

2,599

Deferred revenue - GBT

16,721

145

(4,490

)

12,376

Total contract liabilities

$

22,086

$

145

$

(7,256

)

$

14,975